“yeh ek aise vyakti ki aniyamit ghatna hai jo samay-samay par insaan ko achanak se sadakar, krodhi prani Infernals ke roop mein badal deti hai. Jab Infernals Human Combustion ki pehli pidhi hoti hai, toh dusri aur teesri pidhi pyrokinetics ke roop mein jaani jaati hai – yani aise log jo apne agni ko manipulate aur control kar sakte hain, aur woh insaan bane rehte hain. Infernal khatra ka samna karne aur iska karan jaanne ke liye, Tokyo Armed Forces, Fire Defense Agency, aur Holy Church of Sol ne apna jawab diya: Special Fire Force. Jawan aur utsuk teesri pidhi ke pyrokinetic Shinra Kusakabe, jise apne paon ko ichha ke hisab se jalane ki kshamata se Devil’s Footprints ke naam se jaana jaata hai, woh Special Fire Force Company 8 ke sadasya ban jaate hain. Brigade ki jimmedari ko nibhate hue ki aag mein jal rahe Infernals ko bujha dena aur unki aatmaon ko shaanti dena, Shinra tay hai ki woh wohi hero banega jo agni ke bhay se sankat mein phase hue logon ki jaan bachayega. Lekin yeh woh hero ka khel nahi hai jo Shinra ne socha tha. Fire Force ek toot-phoot ke saath aapas mein jhagadte hue brigades ka ek bikhera hua mess hai, Tokyo ke har kone mein anormal Infernal darshan badh rahe hain, aur ek andheri samuh dawa karta hai ki Shinra ki family ke mrityu ka karan banne wali ajeeb aag ke liye unke paas jawab hai, jo 12 saal pehle hui thi. Fire Force ke andar aur bahar ke kai rukawatein mukabla karne ke liye, Shinra andhere mein use bandhne wale jalan se pichhe ki taraf khoj karta hai.